David P. Penhallow (1854-1910) graduated from the Massachusetts Agricultural College (MAC) in 1873. In 1876, he accompanied W. S. Clark to Japan to help establish Sapporo Agricultural College. He taught botany and chemistry for several years and served as acting president of Sapporo Agricultural College from 1879–80. Upon his return to the United States, Penhallow worked several years at Harvard University with Asa Gray. In 1883, upon Gray’s recommendation he took a position at McGill University, Montreal, Canada. He became a well respected plant physiologist and an expert in fossil conifers. A more complete biography is available at (Dictionary of Canadian Biography).